Aggression (see Glossary) is behaviour proximately intended to cause harm to another individual [1,2], occurring in contexts as diverse as guarding mates, defending territory, competing for nutrients or status, attacking other groups, and protecting or killing infants. From an evolutionary perspective, aggression is expected to protect or increase the aggressor’s fitness on average. Humans are particularly interesting because they pose a behavioural paradox [3,4], simultaneously displaying remarkable tendencies for peace and cooperation [3,5–8] alongside remarkable tendencies for aggression and violence that vary from insults to genocide [3,4,9].

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